Publications

Selected scientific output of the active members of the Center for the Human Past

Jakobsson, M., Bernhardsson, C., McKenna, J., Hollfelder, N., Vicente, M., Edlund, H., Coutinho, A., Sjödin, P., Brink, J., Zipfel, B., Malmström, H., Lombard, M. & Schlebusch, C. (2025). Homo sapiens-specific evolution unveiled by ancient southern African genomes. Nature. https://doi.org/10.1038/s41586-025-09811-4

Schaschl, H., Herzog, T., Oberreiter, V., Kutanan, W., Jakobsson, M., & Larena, M. (2025). HLA diversity and signatures of selection in the Maniq, a nomadic hunter-gatherer population in Southern Thailand. Immunogenetics, 77(1), 23. https://doi.org/10.1007/s00251-025-01380-0

Plascencia, A. G., Jakobsson, M., & Sánchez-Quinto, F. (2025). Ancient DNA HLA typing reveals significant shifts in frequency in Europe since the Neolithic. Scientific Reports, 15(1), 6161. https://doi.org/10.1038/s41598-024-82449-w

de Navascués, M., Burgarella, C., & Jakobsson, M. (2025). Analysis of the abundance of radiocarbon samples as count data. Peer Community Journal, 5. 10.24072/pcjournal.522

McKenna, J., Bernhardsson, C., Waxman, D., Jakobsson, M., Sjödin, P. (2024). Investigating population continuity and ghost admixture among ancient genomes. Hum Popul Genet Genom, 4(3):0009. https://doi.org/10.47248/hpgg2404030009

Breton, G., Barham, L., Mudenda, G., Soodyall, H., Schlebusch, C. M., & Jakobsson, M. (2024). BaTwa populations from Zambia retain ancestry of past hunter-gatherer groups. Nature communications, 15(1), 7307. https://doi.org/10.1038/s41467-024-50733-y

Alves, I., Giemza, J., Blum, M.G.B., Bernhardsson, C., Chatel, S., Karakachoff, M., Saint Pierre, A., Herzig, A.F., Olaso, R., Monteil, M., Gallien, V., Cabot, E., Svensson, E., Bacq, D., Baron, E., Berthelier, C., Besse, C., Blanché, H., Bocher, O., Boland, A., Bonnaud, S., Charpentier, E., Dandine-Roulland, C., Férec, C., Fruchet, C., Lecointe, S., Le Floch, E., Ludwig, T.E., Marenne, G., Meyer, V., Quellery, E., Racimo, F., Rouauld, K., Sandron, F., Schott, J-J., Velo-Suarez, L., Violleau, J., Willerslev, E., Coativy, Y., Jézéquel, M., Le Bris, D., Nicolas, C., Pailler, Y., Goldberg, M., Zins, M., Le Marec, H., Jakobsson, M., Darlu, P., Génin, E., Deleuze, J-F., Redon, R. & Dina, C. (2024). Human genetic structure in Northwest France provides new insights into West European historical demography. Nat Commun 15, 6710. https://doi.org/10.1038/s41467-024-51087-1

Seersholm, F.V., Sjögren, KG., Koelman, J., Blank, M., Svensson, E.M., Staring, J., Fraser, M., Pinotti, T., McColl, H., Gaunitz, C., Ruiz-Bedoya, T., Granehäll, L., Villegas-Ramirez, B., Fischer, A., Price, T.D., Allentoft, M.E., Iversen, A.K.N., Axelsson, T., Ahlström, T., Götherström, A., Storå, J., Kristiansen, K., Willerslev, E., Jakobsson, M., Malmström, H. & Sikora, M. (2024). Repeated plague infections across six generations of Neolithic Farmers. Nature 632, 114–121. https://doi.org/10.1038/s41586-024-07651-2

Bergfeldt, N., Kırdök, E., Oskolkov, N., Mirabello, C., Unneberg, P., Malmström, H., Fraser, M., Sanchez-Quinto, F., Jorgensen, R., Skar, B., Lidén, K., Jakobsson, M., Storå, J. & Götherström, A. (2024). Identification of microbial pathogens in Neolithic Scandinavian humans. Sci Rep 14, 5630. https://doi.org/10.1038/s41598-024-56096-0

Simões, L.G., Peyroteo-Stjerna, R., Marchand, G., Bernhardsson, C., Vialet, A., Chetty, D., Alaçamlı, E., Edlund, H., Bouquin, D., Dina, C., Garmond, N., Günther, T. & Jakobsson, M. (2024). Genomic ancestry and social dynamics of the last hunter-gatherers of Atlantic France, Proc. Natl. Acad. Sci. U.S.A. 121 (10) e2310545121. https://doi.org/10.1073/pnas.2310545121

Zeberg, H., Jakobsson, M. & Pääbo, S. (2024). The genetic changes that shaped Neandertals, Denisovans, and modern humans. Cell 187 (5), 1047-1058. https://doi.org/10.1016/j.cell.2023.12.029

Kırdök, E., Kashuba, N., Damlien, H., Manninen, M.A., Nordqvist, B., Kjellström, A., Jakobsson, M., Lindberg, A.M., Storå, J., Persson, P., Andersson, B., Aravena, A. & Götherström, A. (2024). Metagenomic analysis of Mesolithic chewed pitch reveals poor oral health among Stone Age individuals. Sci Rep 14, 22125. https://doi.org/10.1038/s41598-023-48762-6

Jakobsson, M., Yengo, L., & Zaitlen, N. (2024). Opportunities and challenges for Genomic Data Analyses in Biobanks: a call for papers. Genetics, 226, 4. https://doi.org/10.1093/genetics/iyae023

Bostoen, K., Coutros, P., Doman, J., Fortes-Lima, C., Pacchiarotti, S., & Schlebusch, C. (2025). The Coevolution of Languages, Peoples and Environments in Central Africa’s Kwilu-Kasai Region since~ 1000 BCE: A Dialogue with Jan Vansina. Quaternary Environments and Humans, 100088. https://doi.org/10.1016/j.qeh .2025.100088

Fortes-Lima, C. A., Pacchiarotti, S., Maselli, L., Yugia, J. P. D. B., Kouarata, G., Muluwa, J. K., Mundeke, L., Bostoen, K. & Schlebusch, C. (2025). The Early Bantu Expansion into Central Africa: New Data and Insights from Population Genetics. In An Archaeology of the Bantu Expansion (pp. 172-194). Routledge. DOI:10.4324/9781032658148-9

Lankheet, I., Hammarén, R., Alva Caballero, L. X., Larena, M., Malmström, H., Jolly, C., Soodyall, H., de Jongh, M. & Schlebusch, C. (2025). Wide-scale geographical analysis of genetic ancestry in the South African Coloured population. BMC biology, 23(1), 219. https://doi.org/10.1186/s12915-025-02317-5

Fortes-Lima, C. A., Cerny, V., & Schlebusch, C. M. (2025). Herders from the Green Sahara. American Scientist, 113(4), 232-239. DOI: 10.1511/2025.113.4.232

Bostoen, K., Coutros, P. & Schlebusch, C. (2025). Niger-Congo archaeolinguistics, including Bantu. In: Martine Robbeets, and Mark Hudson (eds), The Oxford Handbook of Archaeology and Language, Oxford Handbooks (2025; online edn, Oxford Academic, 22 July 2025), https://doi.org/10.1093/oxfordhb/9780192868350.013.30.

Fortes-Lima, C. A., Diallo, M. Y., Janoušek, V., Černý, V., & Schlebusch, C. M. (2025). Population history and admixture of the Fulani people from the Sahel. The American Journal of Human Genetics, 112(2), 261-275. https://doi.org/10.1016/j.ajhg.2024.12.015

De Loma, J., Vicente, M., Tirado, N., Ascui, F., Parada, L. A., Gardon, J., Schlebusch, C. & Broberg, K. (2025). Human adaptation in the Andes Mountains. Human Population Genetics and Genomics, 5(1). https://doi.org/10.47248/hpgg2505010002

Larena, M., Chowdhury, A. E., Kels, M. J. T., Tätte, K., Metspalu, M., Schlebusch, C. M., Garcia-Bertrand, R. & Herrera, R. J. (2025). Genetic origins of the Kiritimati population from central-eastern Micronesia. Genome Biology and Evolution, 17(3), evaf046. https://doi.org/10.1093/gbe/evaf046

Steyn, M., Jolly, C., Schlebusch, C. M., & Meyer, A. (2024). Zuurberg (Eastern Cape, South Africa) revisited: human remains, dating and archaeological findings. Azania: Archaeological Research in Africa, 59(2), 294–314. https://doi.org/10.1080/0067270X.2024.2316526

Fortes-Lima, C.A., Burgarella, C., Hammarén, R., Eriksson, A., Vicente, M., Jolly, C., Semo, A., Gunnink, H., Pacchiarotti, S., Mundeke, L., Matonda, I., Koni Muluwa, J., Coutros, P., Nyambe, T.S., Cirhuza Cikomola, J., Coetzee, V., De Castro, M., Ebbesen, P., Delanghe, J., Stoneking, M., Barham, L., Lombard, M., Meyer, A., Steyn, M., Malmström, H., Rocha, J., Soodyall, H., Pakendorf, B., Bostoen, K., & Schlebusch, C.M. (2024). The genetic legacy of the expansion of Bantu-speaking peoples in Africa. Nature 625, 540–547. https://doi.org/10.1038/s41586-023-06770-6

Anderson, C., Scarborough, M., Jocz, L. … Harald Hammarström … & Heggarty, P. (2025). The Indo-European Cognate Relationships dataset. Sci Data 12, 1541 (2025). https://doi.org/10.1038/s41597-025-05445-3

Hammarström, H. (2025). Commentary: Replication, robustness or methodological competition? Linguistic Typology. https://doi.org/10.1515/lingty-2025-0041

Boissonneault, M., & Hammarström, H. (2025). Evaluating the validity of census data for tracking speaker numbers: an investigation of Canada’s Indigenous languages. Current Issues in Language Planning, 1–23. https://doi.org/10.1080/14664208.2025.2524285

Hammarström, H. (2025). On Greenbergs’ classification of African languages. Areas, families, and pools aplenty: a Festschrift for Tom Güldemann. 3-15. https://doi.org/10.18452/32637

Hammarström, H. (2025). Appendices to: On Greenberg’s Classification of African Languages. https://doi.org/10.18452/31664

Elgh, E., Hammarström, H., & Canby, M. (2025). Toward Maximum Parsimony for the Dialect Chain Tree. urn:nbn:se:uu:diva-553592

Hammarström, H., & Borin, L. (2024). A survey of work on Unsupervised Learning of Morphology. Computational Linguistics. https://submissions.cljournal.org/index.php/cljournal/article/view/35

Robbers, M. & Hammarström, H. (2024). Bibliographic bias and information-density sampling. Linguistics Vanguard, 10(1), 239-249. https://doi.org/10.1515/lingvan-2023-0102

Elgh, E. & Hammarström, H. (2024). The dialect chain tree. Diachronica, 41 (3), 307-329.

Forkel, R., & Hammarström, H. (2024). A revised digital edition of Wurm & Hattori’s Language Atlas of the Pacific Area. Sci Data 11, 949. https://doi.org/10.1038/s41597-024-03816-w

Rönchen, P., Wiklund, T., & Hammarström, H. (2024). Likelihood calculation in a multistate model of vocabulary evolution for linguistic dating. Language Dynamics and Change, 14(1), 1-41. https://doi.org/10.1163/22105832-bja10032

Larsson, J. (2025). Exploring the afterlife in Indo-European traditions. In J. Larsson, T. Olander, & A. Jørgensen (Eds.), Indo-European Afterlives: Interdisciplinary Perspectives on Life beyond Death. Stockholm: Stockholm University Press. https://doi.org/10.16993/bcw.a

Larsson, J., Olander, T., & Jørgensen, A. (Eds.) (2025). Indo-European Afterlives: Interdisciplinary Perspectives on Life beyond Death. Stockholm: Stockholm University Press. https://doi.org/10.16993/bcw

Larsson, J., Olander, T. and Jørgensen, A. (eds) (2025) Indo-European Ecologies: Cattle and Milk – Snakes and Water. Stockholm: Stockholm University Press. DOI: https://doi.org/10.16993/bcu.

Larsson, J., Olander, T. & Jørgensen, A.R. (eds.). (2024). Indo-European Interfaces. Integrating Linguistics, Mythology and Archaeology. Stockholm: Stockholm University Press.

Larsson, J. (2024). The many interfaces of Indo-European. In: Jenny Larsson, Thomas Olander & Anders Richardt Jørgensen (eds.) Indo-European Interfaces. Integrating Linguistics, Mythology and Archaeology. Stockholm: Stockholm University Press. 1–2.

Karl-Johan-Lindholm — ^{Karl-Johan Lindholm}

Eriksson, O. & Lindholm, K.-J. (2024). Construction de niche et stratégies des premiers colons dans le centre de la Scandinavie – Écologie historique, pp. 2017.

Arthur, F., Hatlestad, K., Lindholm, K.-J., Loftsgarden, K., Löwenborg, D., Solheim, S., Roche, D. M., & Renssen, H. (2024). The impact of volcanism on Scandinavian climate and human societies during the Holocene: Insights into the Fimbulwinter eruptions (536/540 AD). The Holocene, 34(5), 619-633. https://doi.org/10.1177/09596836231225718

Palmér, A. I. (2025), Indo-Slavic Lexical Isoglosses and the Prehistoric Dispersal of Indo-Iranian. Leiden: Brill. https://brill.com/display/title/72253

Palmér, A. I. (2024). Indo-Slavic lexical isoglosses and the prehistoric dispersal of Indo-Iranian (Doctoral dissertation, Leiden University). https://scholarlypublications.universiteitleiden.nl/handle/1887/3765823

Sjölander, M., Buckland, P. I., & Linderholm, J. (2024). Straight to the point?: Bifacial points and hunter-gatherer mobility in Västerbotten, Sweden. urn:nbn:se:umu:diva-220812

Sjölander, M. (2024). Blinded by the light: developing models of settlement and mobility with the use of spectroscopy and exploratory methods (Doctoral dissertation, Umeå University). urn:nbn:se:umu:diva-220813

Muñoz Mora, V. H., Giannelli, F., Congiu, L., Morales, H. E., & Bertorelle, G. (2025). The hidden threat: genetic load dynamics in tetraploids and diploids. Evolution Letters, qraf036.

Trucchi, E., Massa, P., Giannelli, F., Latrille, T., Gargano, M., Nitta Fernandes, F. A., … & Le Bohec, C. (2025). High gene expression predicts extremely low segregation of deleterious mutations in large penguin populations. Molecular biology and evolution, 42(6), msaf146.

Giannelli, F. (2025). The role of expansion dynamics in shaping genetic diversity and load accumulation: insights from two Lessepsian invaders. (doctoral dissertation, Universitá Politecnica delle Marche) https://iris.univpm.it/handle/11566/342852

Giannelli, F., Obiol, J. F., & Trucchi, E. (2025). Sex-biased dispersal drives mito-nuclear discordance in simulated populations. bioRxiv, 2025-05 (preprint) https://www.biorxiv.org/content/10.1101/2025.05.19.654820v1.abstract

Kouki, R., Giannelli, F., Annabi, K., Dhen, N., Sayadi, A., Essaidi, I., … & Al Mohandes Dridi, B. (2025). DdRAD sequencing and morphometric data analysis reveal a clear differentiation among the Tunisian populations of Onopordum nervosum ssp. platylepis Murb. Genetic Resources and Crop Evolution, 72(4), 4597-4612. DOI: https://doi.org/10.1007/s10722-024-02235-5

Nilsson Stutz, L., Peyroteo Stjerna, R., & Tarlow, S. (2025). Ethical entanglements: human remains, museums and ethics in a European perspective. Antiquity, 99(403), e8. doi:10.15184/aqy.2024.177

Nilsson Stutz, L., Peyroteo Stjerna, R., & Torv, M. (2025). The Oxford Handbook of Mesolithic Europe. urn:nbn:se:lnu:diva-136934

Tõrv, M., & Peyroteo Stjerna, R. (2025). From the Treatment of the Body to the Places of the Dead. In Oxford Handbook of Mesolithic Europe. https://urn.kb.se/resolve?urn=urn:nbn:se:uu:diva-551708

Simões, L.G., Peyroteo-Stjerna, R., Marchand, G., Bernhardsson, C., Vialet, A., Chetty, D., Alaçamlı, E., Edlund, H., Bouquin, D., Dina, C., Garmond, N., Günther, T. & Jakobsson, M. (2024). Genomic ancestry and social dynamics of the last hunter-gatherers of Atlantic France, Proc. Natl. Acad. Sci. U.S.A. 121 (10) e2310545121 https://doi.org/10.1073/pnas.2310545121.

Cveček, S., Herrero-Corral, A., Rebay-Salisbury, K., Banffy, E., Brami, M., Chaix, R., …, Somel. M. … & Žegarac, A. (2025). Beyond Genetics: Exploring Aspects of Non-Biological Kinship in Prehistoric Times. Nature Anthropology, 3(4), 10016.

Yıldız, N., İzgi, H., Rahimova, F., Berkay Altıntaş, U., Gözde Şahin, Z. & Somel, M. (2025). Under the Shadow: Old-biased Genes Are Subject to Weak Purifying Selection at Both the Tissue- and Cell Type-Specific Levels, Genome Biology and Evolution, 17(10), evaf187 https://doi.org/10.1093/gbe/evaf187

Yüncü, E., Doğu, A. K., Kaptan, D., Kılıç, M. S., Mazzucato, C., Güler, M. N., … & Somel, M. (2025). Female lineages and changing kinship patterns in Neolithic Çatalhöyük. Science, 388(6754), eadr2915. DOI:10.1126/science.adr2915

Koptekin, D., Yapar, E., Vural, K. B., Sağlıcan, E., Altınışık, N. E., Malaspinas, A. S., Alkan, C. & Somel, M. (2025). Pre-processing of paleogenomes: Mitigating reference bias and postmortem damage in ancient genome data. Genome biology, 26(1), 6. https://doi.org/10.1186/s13059-024-03462-w

Koptekin, D., Aydoğan, A., Karamurat, C., Altınışık, N. E., Vural, K. B., Kazancı, D. D., … & Somel, M. (2025). Out-of-Anatolia: cultural and genetic interactions during the Neolithic expansion in the Aegean. Science, 388(6754), eadr3326. https://doi.org/10.1126/science.adr3326

Mazzucato, C., Coscia, M., Küçükakdağ Doğu, A., Haddow, S., Kılıç, M. S., Yüncü, E., & Somel, M. (2025). A Network of Mutualities of Being: Socio-material Archaeological Networks and Biological Ties at Çatalhöyük. Journal of Archaeological Method and Theory, 32(1), 25. https://doi.org/10.1007/s10816-024-09692-3

Szeniczey, T., Melis, E., Jakab, J., Bátora, J., Major, I., Molnár, M., … Reyes Centeno, H., … & Hajdu, T. (2025). The population history of Central Europe in the Early Bronze Age (2200/2100–1600/1500 BC)-a craniometric approach. Archaeological and Anthropological Sciences, 17(11), 1-21. DOI: https://doi.org/10.1007/s12520-025-02324-1

Adel, M., Hunt, K. J., Lau, D., Hartsfield, J. K., Reyes-Centeno, H., Beeman, C. S., Elshebiny, T., & Sharab, L. (2025). Precision Assessment of Facial Asymmetry Using 3D Imaging and Artificial Intelligence. Journal of Clinical Medicine, 14(20), 7172. https://doi.org/10.3390/jcm14207172

Keppeler, J. G., Rodriguez, M., Koontz, S., Wise, A., Mink, P., Crothers, G., Murphy, P. R., Robertson, J. K., Reyes-Centeno, H., & Uhl, A. (2025). Remote Sensing of American Revolutionary War Fortification at Butts Hill (Portsmouth, Rhode Island). Heritage, 8(10), 430. https://doi.org/10.3390/heritage8100430

Benitez, D., Delgado, M., & Reyes-Centeno, H. (2025, March). The application of nonmetric dental traits in analyzing sex mobility and social networks among pre-Hispanic societies from the Northern Andes. In AMERICAN JOURNAL OF BIOLOGICAL ANTHROPOLOGY (Vol. 186, pp. 14-15). 111 RIVER ST, HOBOKEN 07030-5774, NJ USA: WILEY.

Jakob, A. (2025). Studies in East Baltic consonant clusters 1:* šr. Baltistica, 60(1), 7-29. DOI: 10.15388/baltistica.60.1.2578

Jakob, A. (2025). Studies in East Baltic consonant clusters 2:* rTv. Baltistica, 60(2), 241-256. DOI: 388/baltistica.60.2.2581

Yamazaki, Y. (2023). The simple thematic present ending in the 2nd person singular in Baltic. The Journal of Indo-European Studies, 51(1&2), 147-164.
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